What constitutes a major threat to the survival of turtles a

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What constitutes a major threat to the survival of turtles according to Elizabeth Griffin?
此句的主语是what吗

What constitutes a major threat to the survival of turtles according to Elizabeth Griffin?
根据伊丽莎白·格里芬,什么对海龟的继续生存构成了主要威胁?
疑问词 what 就是这句话的主语,谓语动词是 constitute,其后的 a major threat 是宾语.

What constitutes a major threat to the survival of turtles a 英语翻译请翻译：The defeat constitutes a major set-back for our dipl The survival of civilization as we know it is ___ threat. THE JEWISH PEOPLE A STORY OF SURVIVAL怎么样 Overpopulation poses a terrible threat to the human race.Yet Is rising China a 'threat' to the world debate:does human civilization become a threat to the earth? the principle of survival of the fittest is one of the major .What is a federal system? What are some of the major differ what a major bitch 1,ISall species of sea turtles are endangered because （the a 用pose a great threat to造句